SICB Division of Invertebrate Zoology (DIZ)

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From the DIZ Researchers Database

Svetlana Maslakova

Evolution of Larval Development in Nemerteans
Nemerteans (Phylum Nemertea) belong to the protostome clade the Trochozoa, which includes annelids, mollusks, sipunculids, echiurids and entoprocts. All trochozoan phyla, except nemerteans, have been shown to possess a trochophore larva characterized by the prototroch, a pre-oral belt of specialized ciliated cells derived from the specific founder cells called the trochoblasts. Prototroch is the primary locomotory organ of many larvae and is also often involved in feeding. Nemerteans possess a wide diversity of larval body plans, but until now no nemertean had been shown to possess a trochophore larva or a prototroch, which hampers comparisons between the larval development of nemerteans and other trochozoans. Members of the nemertean clade Pilidiophora develop via a highly specialized planktonic pilidium larva, which undergoes catastrophic metamorphosis, while members of the sister clade to Pilidiophora - the order Hoplonemertea, and the basal palaeonemerteans have uniformly ciliated "planuliform" larvae, which develop into juvenile without a drastic change of the body plan. The answer to which type of development - with or without pilidium - is primitive for the nemerteans has implications for the evolution and loss of novel complex larval forms (such as pilidium). In collaboration with Mark Martindale (University of Hawaii) and Jon Norenburg (Smithsonian Institution) we used intracellular labeling markers, cLSM and SEM to demonstrate that uniformly ciliated larva of a palaeonemertean Carinoma tremaphoros possesses a hidden prototroch which is derived from the same cell lineage as prototrochs of other Trochozoans. This finding refutes the homology between the pilidium and the so-called larval ectoderm previously reported from some palaeonemertean larvae – the main support for idea that pilidial development is ancestral for the phylum and instead suggests that pilidium is an elaboration on the trochophore groundplan, uniquely derived within nemerteans. Additionally, the inferred ancestral presence of a trochophore in nemerteans fills the gap in the distribution of trochophore larvae among the Trochozoa and allows a meaningful comparison between larval development of nemerteans and other trochozoans.


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Officers & Representatives


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Chair 01/2009-01/2012 Dianna K. Padilla
Past Chair 01/2009-01/2011 Janice Voltzow
Secretary 01/2007-01/2010 Renae Brodie
Program Officer 01/2008-01/2011 James B. McClintock
Student/Postdoc Rep. 01/2008-01/2011 Jann Vendetti
Libbie Hyman Memorial Scholarship Committee Chair Bruno Pernet




DIZ Memberlist

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Tara   University of Arkansas 
Peter James Adam  Northwest Missouri State Univ 
Roger Adams   
Nikki L. Adams  CA Polytechnic U 
Christina Aimone   
Nicholas Alcorn  Bowdoin College 
David E. Alexander  University of Kansas Department of EEB 
Jonathan D. Allen  Randolph-Macon College 
Brett Allen  Blackburn College 
Pamela Ander  Brooklyn College 
John M. Anderson   
David Anderson   
Anthony T. Annunziato  Boston College 
Jorg Anson  University of Hawaii 
Tomoji Aoto   
Cesar Arenas-Mena  San Diego State 
Emily Arnold   
Audrey Aronowsky  The Field Museum 
Akira Asakura  Nat Hist Mus Chiba 
Kyle Aveni-Deforge   
Lisa Marie Axtman  Colorado State Univ 
Gregory Alan Babbitt  Arizona State University 
Shirley M. Baker  University of Florida 
Elizabeth J. (Susie) Balser  Illinois Wesleyan U 
Michael Baltzley  St Mary's College of Maryland 
Flavia Barbosa  University of Missouri 
Penelope A. G. Barnes  Malaspina Univ 
Franklin H. Barnwell  University of Minnesota Dept of Ecology Evolution & Behavior 
John Barthell  Univ of Central Oklahoma 
Ian Bartol  Old Dominion University 
Lawrence V. Basch  University of Hawaii 
Anna Lee Bass  Univ of New England 
Raymond Bauer  Univ Louisiana 
Christopher J. Bayne  Oregon State University 
Richard David Beckwitt  Framingham State College 
Mark A. Beekey  Sacred Heart Univ 
Tina Marie Bell  University of Georgia 
Jennifer Bell  UC Berkeley 
Gerard L. Bellan  Centre National de La Recherche Scientifique 
Miri Ben-Hamo   
Rebecca Beresic-Perrins  N Arizon Univ 
Sarah Kosting Berke  Smithsonian--SERC 
Jennifer Bernatis  Univ. of Florida 
Elena Bernick  WA State Univ 
Patricia M. Biesiot  U of S Mississippi 
Ruediger Birenheide   
Cory Bishop  Dalhousie Univ 
Craig Black   
Todd Blackledge  U Akron 
Neil W. Blackstone  Northern Illinois University 
Sydella Blatch  Arizona State 
Cherissre Boateng  UVI 
Stefanie Anne Boettger  West Chester Univ 
Michael Boller  Saint John Fisher College 
Calhoun Bond  Greensboro College 
Charles E. Booth  Eastern Connecticut State Univ 
Elizabeth Borda  Auburn University 
Stephen Borgianini  University of South Carolina Beaufort 
Isidro Bosch  SUNY at Geneseo 
Jason T Botz  KU 
George Bernard Bourne  Univ of Calgary 
Milton J. Boyd  Humboldt State University 
Barbara C. Boyer  Union College 
Sarah L Boyer  Macalester College 
Sean Brady  Smithsonian 
Claire M Brett  Univ of St Andrews 
Robert H. Brewer  Trinity College 
Ashley Kristen Brewington   
Kevin Brix  University of Miami 
H. Jane Brockmann  University of Florida Dept of Zoology 
Renae Brodie  Mount Holyoke College 
Stine Brown  University of New England Department of Life Sciences 
Federico Brown  Max Planck Inst for Dev Biology 
Rena Bryan  Univ of Florida 
Roger A Bryne  State University of New York 
David Buchwalter  North Carolina State University 
Graham Budd  Uppsala University 
Jennifer Burnaford  California State University Fullerton 
Louis E. Burnett  College of Charleston 
Andrew Butz  University of Cincinnati Department of Biological Sciences 
Roger A. Byrne  SUNY Fredonia 
Maria Byrne  Univ of Sydney 
Candice Bywater  University of QLD 
Roy L Caldwell  Univ of CA Berkeley 
Chris B Cameron  University de Montreal 
Terry G. Campbell  Univ of South Florida 
Eric Octavio Campos   
Johanna Cannon  Auburn University 
Manuela Caprioli  Univ of Milan 
Angela Caranci   
Jason Carbaugh  SE Missouri State U 
Paulyn Cartwright  University Kansas 
Jodi Caskey  U of LA-Lafayette 
Frank L. Castille  TAES Shrimp Mariculture Research 
Cynthia Catton  UCSD 
Kimberly Catton  Georgia Tech 
Michael J. Cavey  University of Calgary 
Shella Cervantes  CalAcademy 
Nanette E. Chadwick  Auburn University 
Carol Lynn Chaffee  Univ of Florida 
Ernest S. Chang  University of California 
Kourt Chatelain  Weber State Univ 
Ming Chen  U Conn 
Aaron Cheng   
Alex G. Cheroske  University of Maryland Baltimore County 
Alexander Cheroske  Mesa Community College 
Kai-Jung Chi  Nat. Chung Hsing Univ., Taiwan 
Ariel Chipman  The Hebrew Univ of Jerusalem 
Il-Gyu Cho   
Katie Chronister   
Ka Hou Chu  The Chinese University of Hong Kong 
John C. Clamp  North Carolina Central University 
Martine B. Claremont  NHM, London 
Mary E. Clark   
Andrew Clark  UC Irvine 
Kathryn A. Coates  Bermuda Dept of Consv Services 
Melissa Coates  Univ of CA, Los Angeles 
Mary Alice Coffroth  SUNY at Buffalo 
C. Sarah Cohen  Romberg Tiburon Center, SFSU 
Jonathan Cohen  Harbor Branch Oceano. Inst. 
Itai Cohen  Cornell University 
Sean P Colin  Roger Williams Univ 
Rachel Collin  STRI 
Allen G. Collins  Smithsonian Inst, MRC 0163 
Stacey Combes  Harvard University 
Thomas R Consi  Univ of Wisconsin-Milwaukee 
Heidy Contreras   
Clayton B. Cook  Harbor Branch Oceanographic 
Susan B. Cook  Consortium for Ocean Leadership 
Steven L. Coon  National Institutes of Health 
Marla Coppolino  Southern Illinios University 
John O. Corliss   
John C. Cornell   
Raena M Cota  N Arizona Univ 
Jonathan D Cowart  UNC Wilmington 
David L. Cowles  Walla Walla College 
Nicole Cox  Clemson University 
Jonathan D Craft  College of Charleston 
Bruce J. Crawford  University of British Columbia 
Sherine Crawford  Medgar Evers?CUNY 
Stephanie Crofts  UC Irvine 
Roger Croll  Dalhousie Univ 
Peter James Croucher  ESM, Univ CA Berkeley 
Andrea Louise Crowther   
Lisa Crummett  UFL 
Lawrence A. Curtis  University of Delaware Cape Henlopen Laboratory 
Nicholas Edward Curtis  USF 
Dan Curtis  UNLV 
Paul A Cziko  U of OR 
John O Dabiri  California Institute of Technology 
Thomas L. Daniel  University of Washington Department of Zoology 
Melissa Daugherty  Cal Poly SLO 
Ian Robert Davenport  Xavier Univ of Louisiana 
Katie Davenport  Swarthmore College 
Goggy Davidowitz  University of Arizona 
Brad Davidson  Univ of Ariz 
Kiyya Davis  Medgar Evers 
Elizabeth Davis-Berg  Columbia College Chicago 
John H. Dearborn   
Lewis E. Deaton  Univ of Louisiana at Lafayette 
Sandie M Degnan  University of Queensland 
Marta J. DeMaintenon  U Hawaii at Hilo 
J. Manuel Denuce  Radboud University 
Charles Derby  Georgia State University 
Shahan Derkarabetian   
Maya deVries  UC Berkeley 
Ruth Ann Dewel   
William C. Dewel  Appalachuan State University 
Gary Howard Dickinson  Duke Marine Lab 
Jennifer Dijkstra  Wells National Estuarine 
Michael Dillon  UC Berkeley 
Ronald V. Dimock  Wake Forest University 
Jan-Henning Dirks  University of Cambridge 
William E. Dobson   
David Doe  Westfield State College 
Seth Donoughe  Swarthmore College 
Thomas Michael Dornhoffer   
John K Douglass  Smithsonian Tropical Research Institute 
William L. Downing  Hamline University 
Michelle D Drewry  Univ of North Florida 
Christopher Anson Drummond  Bowling Green State Unive 
Benjamin Dubansky  LSU 
Jeff Dudycha  Univ of South Carolina 
Casey W Dunn  EEB Brown University 
Mike Dunn  Blackburn College 
Paul Richard Dunn   
Kevin J. Eckelbarger  University of Maine 
Sarah Edmonds  Univ Akron 
Dale Edwards  Univ of Evansville 
Olaf Ellers  Free Lance 
Joel K. Elliott  University of Puget Sound Department of Biology 
Glen Elliott  University of Alberta 
Richard B. Emlet  Univ Oregon 
Patrick Erwin  U Alabama at Bham 
Anne M. Estes  University of Arizona 
Shelley Etnier  Butler Univeristy 
Peter Etnoyer  Harte Research Institute 
Dennis J Evangelista  UC Berkeley 
Jan Robert Factor  Purchase College, SUNY 
Roger Dean Farley  University of California 
Anuschka Faucci  Univ of Hawaii 
Zen Faulkes  Univ of Texas-Pan American 
Daphne Gail Fautin  University of Kansas 
Horst Felbeck  University of California San Diego 
William E. Fennel   
John C. Ferguson  Eckerd College 
Daniel Fernandes   
Joan D. Ferraris  NIH/NHLBI/LKEM 
David L Ferrell  Florida State University 
David Ferrier  Univeristy of St Andrews 
Jeremy Fields  University of San Diego 
John R. Finnerty  Boston University 
Larklyn H. Fisher   
Desiree Fleming  Blackburn College 
Megan Flenniken  Stony Brook University 
Peter P. Fong  Gettysburg College 
David Forest  UMIT 
Juan Jose Fraire-Zamora  University of California Riverside 
Tamara Frank  Harbor Branch Oceanographic 
Markus Frederich  Univ New England 
Andrea Frey  UNH 
Robert Full  UC of California at Berkeley 
Peter Funch  Univ of Aarhus 
Megumi M Fuse  San Francisco State University 
Jonathan Gain   
Louis F. Gainey  Univ Southern ME 
Stephen L. Gardiner  Bryn Mawr College 
Yvette Garner  UNH 
Jonathan B. Geller  MLML 
Sophie George  Georgia Southern University Department of Biology 
Gretchen Gerrish   
Michael T. Ghiselin   
Victoria Gibbs  Univ of AL at Bham 
Cole Gilbert  Cole University 
Sandra L. Gilchrist  New College of Florida 
Rhanor Gillette  University of Illinois 
Casey Gilman  UNM 
David W Ginsburg  Univ of California, Los Angeles 
Peter R Girguis  Harvard Univ 
Jeffrey Goldman  Center for Embedded Network Sensing 
Andrea Gomez  NYU School of Medicine 
Jesus Gomez  UPR Humacao Campus 
Vanessa Gonzales  CSU Fresno 
John Stephen Gosnell  University of Californa, Santa Barbara 
Annette Govindarajan  Woods Hole Oceanographic Institute 
Robert Grabowski  Queen Mary, University of London 
Robert A Graham  Rubino Academy 
Judith P. Grassle  Rutgers State Univ of NJ 
Frank William Grasso  Brooklyn College 
Sarah Amelia Gravem  California Polytechnic State Univ 
Jonathan Green  Roosevelt University Department of Biology 
Mike Greene  CU-Denver 
Brian Gregory   
Daniel Grunbaum   
Daniele Guinot  MNHN, Paris 
Gilbert F. Gwilliam  Reed College Department of Biology 
Steve Haddock  MBARI 
Michael G. Hadfield  Univ of Hawaii 
Stavros P Hadjisolomou  Brooklyn College 
Steve Hageman  Appalachian State 
Kenneth M. Halanych  Life Sciences Dept 
Christina Hamlet  Univ of North Carolina 
Jorg U Hammel  FSU Jena Germany 
Carl S. Hammen   
Steven C. Hand  Louisiana State Univ 
Shanna Dawn Hanes  Auburn University 
John P. Harley  Eastern Kentucky University 
Larry G. Harris  University of New Hampshire 
Shaunna Harris  Louisiana State Univ 
Frederick W. Harrison  Journal of Morphology 
Michael W. Hart  Simon Fraser University 
Alexandra Hart  University of Washington 
Scott D. Hawke  Willamette University 
Kenneth A. Hayes  Smithsonian Inst Nat'l Mus of Nat Hist 
Hilary A Hayford  Moss Landing Marine Labs 
Jennifer Head  OIMB 
Elizabeth Anne Chapman Heath-Heckman  Univ of Wisconsin-Madison 
Tyson Hedrick  UNC at Chapel Hill 
Esptit Heestand  The Ohio State University 
Thomas Hegna   
Andreas Hejnol  University of Hawaii 
Bryan Robert Helm  Univ of Arizona 
Brian Helmuth  University of South Carolina 
Gordon Hendler  Natural History Museum Research and Collections 
Raymond P. Henry  Auburn University 
Henriette Herlant-Meewis   
Colin Olmsted Hermans  Sonoma State University 
Pablo JAVIER Hernandez-Garcia  Institute for Tropical Ecosystem Studies 
Jane Hey  Morningside College 
Joseph K. Hichar   
Carole S. Hickman  UC Berkeley 
Michio Hidaka  University of the Ryukyus Department of Biology 
Robert P. Higgins   
John G. Hildebrand  University of Arizona ARL Division of Neurobiology 
Robert B. Hill  University of Rhode Island Department of Biological Sciences 
Ryan Isaac Hill  UC Berkeley 
Susan Hill  Michigan State University 
Anson H. Hines  Smithsonian Environmental Research Center 
Kenneth David Hoadley  Univ of North Carolina @ Wilmington 
Jeffrey M Hoch  SUNY-Stony Brook 
Rick Hochberg  University of Massachussetts Lowell 
Edward S. Hodgson   
Jason Hodin  Hopkins Marine Station 
Cheryl Hogue  California State Univ, Northridge 
Charles W. Holliday  Lafayette College 
Farland Holliman  NC State University 
Eric R. Holm  NSWC Carderock 
Alan Holyoak  BYU-Idaho Dept of Biology 
Thomas Honegger  University of Zurich 
Matthew Hooge  Univ of Maine 
Mildred A Hoover  Salem State College 
Melanie Hopkins  Univ of Chicago 
Francis Ray Horne  Texas State University 
Hsin-Drow Huang  Academia Sinica 
Ying Huang   
Christine Huffard  UC Berkeley 
Turkesha Huggins  Medgar Evers 
William D. Hummon  Ohio University 
Marguerite Hunt  Univ Texas, Austin 
Sofia Malinda Hussain  Univ of S Florida 
Fumio Iwata  Laboratory of Nemertea 
Erika V. Iyengar  Muhlenberg College 
Jennifer Jackson  Georgia Tech 
Molly W. Jacobs  U Connecticut 
David Jacobs  UCLA 
Claire James  Univ of Portland 
Jason E Jannot  Purdue University 
William B. Jeffries  Dickinson College 
John Jellies  Western Michigan University 
Ronald Adam Jenner  University of Bath 
David Jennings  Univ of South Florida 
Holly Jessop  U of Hawaii at Manoa 
Houshuo Jiang  Woods Hole Oceanographic Institution 
Elisha Jno-Baptiste  Univ of Virgin Islands 
Sonke Johnsen  Duke University 
Amy S Johnson  Bowdoin College 
Kevin Johnson  Florida Institute of Tech 
Sheri L Johnson  University of Florida 
Collin Johnson  Harvard University 
Rebecca Fay Johnson  California Academy of Sciences 
Barbara Joos  Sable Systems 
Karl H Joplin  East Tennessee St Univ 
Jennifer Jost  USC Columbia 
Jennifer Jost  U of New England 
Timothy Judd  SE Missouri State Univ 
Peter A. Jumars  University of Maine Darling Marine Center 
Steven H. Jury  Marical 
Robin W. Justice  The Claremont Colleges 
Suzanne Kacenas  Col of Charleston 
Isabella Kappner  American Museum of Natural History 
David Karlen  Environmental Protection Commission of Hillsborough County 
Masaya Katoh  Seikai National Fisheries Research Institute Ishigaki Tropical Station 
Susan L. Keen  UC-Davis 
Stephen C. Kempf  Auburn University 
William M. Kier  UNC - Chapel Hill 
Takahiro Kikawada  NIAS Japan 
Alison Jane King  Dalhousie University 
Candice King  Medgar Evers 
Roni Kingsley   
Stephen T. Kinsey  UNC Wilmington 
Lisa A. Kirkendale  Univ of Florida 
Thomas Scott Klinger  Bloomsburg University 
Emily Knott  Univ of Jyvaskyla 
Robert E. Knowlton  GWU 
Markus Koch  Freie Universitaet of Berlin 
Kevin Kocot  Auburn University 
Mimi A. R. Koehl  UC Berkeley 
Joris M Koene  Vrije Universiteit 
Alan J. Kohn  University of Washington 
Talina Konotchick  Scripps Institution of Oceanography 
Gil Koplovitz  UAB 
Ellen Kosman   
Jeffrey J Kovatch  Marshall University 
Sindhu Krishnankutty   
Janice L Krumm  Widener University 
Joseph G. Kunkel  University of Mass - Amherst 
Dian-Han Kuo  UC Berkeley 
Lena Kupriyanova  U of Adelaide 
Samantha Kushner   
Lisa Kvalheim   
Michael LaBarbera  University of Chicago Organismal Biology & Anatomy 
Rene LaFont  Universite Pierre et Marie Curie 
Harry H. Laidlaw  University of California, Davis Department of Entomology 
Matthew P. Landau  Richard Stockton College 
Thomas Langley  UNH 
Claire Larroux  U of Queensland 
Howard R. Lasker  University at Buffalo 
Christopher Laumer  Harvard University 
Dennis Lavrov   
Ronald G. Lawler   
John M. Lawrence  University of South Florida 
Hilary Lease  Univ of New Mexico 
Sung Gu Lee  Colorado State Univ. 
Jennifer Legere  Dalhousie Univ 
Lucy Leid  Medgar Evers 
Esther M. Leise  UNCG Dept of Biology 
Howard M. Lenhoff   
Michael P. Lesser  University of New Hampshire 
Herbert W. Levi  Harvard University Museum of Comparative Zoology 
Lisa Levin  Scripps Inst Ocean 
Don Richard Levitan  Florida State Univ 
Sylvia Lewis  C of C 
Sally Leys  University of Alberta 
Bruce Lieberman  University of Kansas 
Hui-Chen Lin  Tunghai University Department of BIology 
Huai-Ti Lin  Tufts University 
David Richard Lincicome   
David R. Lindberg  University of California 
Annie Lindgren  The Ohio State University 
Sara M. Lindsay  U Maine 
Kristin Lobo  St Josephs Univ 
John H. Lochhead   
Jan M. Locke   
Tse-Lynn Loh  University of North Carolina - Wilmington 
Stephen H. Loomis  Connecticut College 
Paola Lopez-Duarte  Scripps Inst Ocn 
Tito Monteiro de Cruz Lotufo  Universidade Federal do Ceara 
Eric C Lovely  Arkansas Tech University 
Donald L. Lovett  College New Jersey 
Lara Luzak  Hofstra University 
Tara Magi  Walla Walla Univ 
Roxane Magnus   
Christopher L. Mah  National Museum of Natural History 
Andrew R Mahon  University of Notre Dame 
Heidi Mahon  Old Dominion Univ 
Elisa Maldonado  UCSD - Scripps Inst of Oceanography 
Max Maliska  Univ of Washington 
Arun Mani   
Linda H. Mantel   
Robert C. Maris  Mansfield University of Pennsylvania 
David B Mark Welch  Marine Biological Laboratory 
Daniel Marshalonis  Dept. of Biological Sciences 
Gary G. Martin  Occidental College 
Kesha Martin  Medgars Evers College 
Joel Martin  NHMLAC 
Vicki J. Martin  Appalacian State Univ 
Mark Q. Martindale  Univ of Hawaii 
Gabriela Martinez  U of New Hampshire 
Svetlana Maslakova  University of Oregon 
David Matus  Univ of Hawaii 
Mikhail Matz  UT Austin 
Tim Maugel  University of Maryland 
Amy B Maxmen  Harvard University 
Maria Mazzillo  Auburn University 
Justin McAlister  Clemson University 
William McBlair   
Michael A McCartney  UNC Wilmington 
Marion McClary  Fairleigh Dickinson 
James B. McClintock  UAB 
John McConaugha  Old Dominion University 
Adam M McCoy  Hopkins Marine Station 
Kwanza McCoy  Medgar Evers College 
John J. McDermott  Franklin and Marsh College 
Kathryn A McDonald  Smithsonian Tropical Rsrch Inst 
Catherine S. McFadden  Harvey Mudd College 
Wendy McFarlane  Manhattanville College 
Iain James McGaw  Univ of Nevada Las Vegas 
Nature McGinn  Univ of California, Davis 
Tamara M. McGovern  Clemson University 
Matthew J McHenry  U. C. Irvine 
Damhnait McHugh  Colgate University 
Ellen W. McLaughlin  Samford University 
Robert F. McMahon  U Texas Arlington 
Kristina S. Mead  Denison University 
Matthew J Medeiros  UC Berkeley 
Lorenzo Menzel  Florida International University 
Neil A. Mercando  Penn State University Abington College 
Sara Merlie  Winthrop U. 
Rachel Ann Merz  Swarthmore College 
Erin Leigh Meyer  Univ of California Berkeley 
Eli Meyer  University of Texas at Austin 
John Meyer  Univ of NH 
Constantine I Michaelidis  Swathmore College 
Francois Michonneau  University of Florida 
Maria-Pia Miglietta  Pennslyvania State 
Cecelia Miles  University of Chicago 
E. Morton Miller   
Stephen A. Miller  College of the Ozarks 
Laura Miller  University of North Carolina-Chapel Hill 
Luke Paul Miller  Northeastern Univ 
Nathan Miller  Albion College 
Wyatt Austin Miller   
Abraham Miller  Univ of Tampa 
Alessandro Minelli  University of Padova 
Benjamin Gregory Miner  Western Washington University 
Patricia Mire  UL Lafayette 
Michael J. Misamore  Texas Christian U 
Brittany Misercola  St Lawrence Univ 
Toni Mizerek  San Diego State University 
Ortal Mizrahy   
Andrew Mobley  UAB 
Stacia Moffett  Washington State University 
Yamile Molina   
Emilee Monson  UAA 
Richard Mooi  California Academy of Sciences 
Anne M. Moore  Univ of the Pacific 
Amy Moran  Clemson Univ. 
Cheryl Morrison  USGS Leetown WV 
John Morse  UT-Arlington 
M. Patricia Morse   
Tatsuo Motokawa  Tokyo Institute of Technology Basic Biology 
Jennifer Mraz  Univeristy of Southern Mississippi 
Nancy Muehllehner  Cal State Univ N 
Charles Mulligan  University of Florida 
Marie Myrthil  Medgar Evers College 
Lisa M. Nagy  University of Arizona Dept. of Molecular & Cellular Biology 
Annalise Nawrocki  University of Kansas 
Brian T. Nedved  Univ of Hawaii Dept Zoology/PBRC 
Matthew Nelson  NIWA 
Michael Nelson  Medgar Evers/CUNY 
Michael S. Newel  University of Calgary 
Peter Ng  NUS 
Scott Nichols  University of California 
Michael Nickel  Institut fuer Spezielle Zoologie und Evolutionsbiologie, FSU Jena 
Michele Kiyoko Nishiguchi  New Mexico State University 
Marie-Therese Noedl   
Suegene Noh   
Jon L. Norenburg  Smithsonian Institution 
Albert Charles Nyack   
James W. Nybakken  Moss Landing Marine Laboratories 
Spencer Nyholm  University of Connecticut 
Micha J O'Donne   
Michael O'Donnell  Friday Harbor Laboratories 
Brian O'Meara  NESCent 
Beth Okamura  NATURAL HISTORY MUSEUM 
Robert K. Okazaki  Weber State Univ 
AdeGbenga Adekunle Okuwoga  Epetedo senior High School 
Tyler J Olivier  Univ of Louisiana Lafayette 
Horst Onken  Washington State 
Brian Ortman  UNH 
Fernanda Ximena Oyarzun  U of Washington 
Dianna K. Padilla  Stony Brook University 
Louise R. Page  Univ of Victoria 
Emma Palacios Theil  ULL 
Megan Partyka  Tulane University 
Sheila Patek  UC Berkeley 
Darshan Patel  Swarthmore College 
Steven Patten  Walla Walla Univ 
John S. Pearse  University of California Santa Cruz Institute of Marine Sciences 
Vicki Pearse   
Jan A. Pechenik  Tufts University 
Nancy Pelaez  Purdue University Bio Sci 
Peltin Pelep  UH at Manoa 
Chuck Pell  Duke University 
Andrew Pemberton  Univ of Zurich 
D. Carolina Penalva-Arana  Indiana University 
Yamel Perdomo  Medgar Evers 
Santiago F. Perez  Stanford University 
Kaipo Perez III   
Omar Perez Reyes  University of Puerto Rico 
Marcos Perez-Losada  BYU 
Bruno Pernet  California State Univ/Long Beach 
Elizabeth Perotti  UC Berkeley 
Emily Petchler  Muhlenberg College 
Kevin Peters  UAB 
Mary Elizabeth Petersen  Darling Marine Center 
Janne Pfeiffenberger  USF 
Sujal Phadke  University of Houston 
Nicole E. Phillips  Victoria University of Wellington 
Sidney K. Pierce  Univ South Florida 
John F. Pilger  Agnes Scott College 
Tia Renee Pilikian   
Anthony Pires  Dickinson College 
Marina Piscitelli   
David Plachetzki  UCSB 
Robert D. Podolsky  College of Charleston 
Lisa Portis  Colby College 
Kristen Ann Potter  Univ of Arizona 
Mickie L. Powell  UAB 
Marney C. Pratt  Mt Holyoke College 
Robert L. Preston  Illinois State University 
Robert S. Prezant  Montclair State University 
Rebecca Price  UNC Chapel Hill 
Austin W. Pritchard  Oregon State University Department of Zoology 
Dina Proestou  Univ of Rhode Island 
Rachel Przeslawski  Stony Brook University 
Sakshi Puri  Univ of Texas -Pan American 
Hollie Putnam  Univ of Hawaii 
Michael C. Quinlan  Midwestern University Department of Physiology 
Brenda Rascon  Arizona State Univ 
Maggie Ray  NC State University 
Abigail Reft   
Carl L. Reiber  UNLV 
Chris Reinhard  Univ of Kansas 
Kathleen Reinsel  Wittenberg University 
Xiaoyun Ren  UC Berkeley 
Patrick D. Reynolds  Hamilton College 
Mary E. Rice  Smithsonian MarSta 
Michael A. Rice  University of Rhode Island 
Stanley A. Rice  University of Tampa 
Ammon Rice  CSU Fresno 
Earl R. Rich  University of Miami Department of Biology 
Jeff Riffell  UCLA 
Marc Daniel Rinehart   
Brian Rivest  SUNY at Cortland 
John L. Roberts  University of Massachusetts, Amherst Department of Biology 
Larry S. Roberts  Florida International Univ 
William E. Robinson  University of Massachusetts, Boston 
Eve Robinson  Univ of California, Berkeley 
Morris Rockstein   
Maurico Rodriguez-Lanetty  Oregon State University 
Pamela Roe  California State University, Stanislaus 
Constance Lynn Rogers-Lowery  Catawba College 
Sandra L. Romano  University of the Virgin Islands 
Jocelyn Romano  Duke University 
Melissa Romero  CSU, Los Angeles 
Rebeca Rosengaus  Northeastern Univ 
Louise Russert-Kraemer  University of Arkansas 
Kevin Alexander Ryan  Winthrop University 
Nathan Rycroft  Old Dominion Univ 
Krystal Rypien  Scripps Institution of Oceanography 
Mary Beth Saffo  Harvard University 
Kenneth S. Saladin  Georgia College and State Univ. Dept. of Biological & Environ. Sci. 
Dahniel Samuel  Medgar Evers/CUNY 
Allen F. Sanborn  Barry University 
Matthew Sanders  CEFAS 
Thomas Sandor   
David Sandstrom  National Institute of Mental Health 
Scott Santagata  C.W. Post Campus of Long Island University 
Noriyuki Satoh  Kyoto University Department of Zoology 
Sara J. Sawyer  Glenville State College 
John Scarpa  Harbor Branch Oceanographic Institute at Florida Atlantic 
Lindsay Schamel  St Joseph's Univ 
John I. Scheide  Central Michigan University 
Rudolf S. Scheltema  Woods Hole Oceanographic Institution 
Steve Contos Schenk  Truckee Meadows Community College 
Bernd Schierwater  ITZ Hannover 
Frederick R. Schram  Burke Museum 
Paul C. Schroeder  Washington State University 
Nancy Schult  Colgate Univ 
Anja Schulze  Smithsonian 
Megan Schwartz  Seattle University 
Julie Schwartz  U of S Florida 
Ryan Schwarz  Univ of New Mexico 
Kenneth P. Sebens  University of Washington 
Andrew Sensenig   
Todd Sformo  Univ of Alaska Fairbanks 
Asaf Shechter  Ben Gurion Univ 
Kristin M Sherrard  Friday Harbor Laboratories 
J. Malcolm Shick  University of Maine School of Marine Sciences 
Chris Shields  Clemson University 
Nicholas Shikuma  Univ of Hawaii 
Thomas C. Shirley  Harte Research Institute 
Stanley Shostak  University of Pittsburgh 
Shigeno Shuichi  University of Chicago 
Sandra E. Shumway  Uconn 
Julia Sigwart  Univ College Dublin 
James Sikes  Univ of Maryland 
Joana Zanol Silva  George Washington University 
Michael Simon  Tufts University 
Andrew Simpson  UC Riverside Paleo 
Anne Simpson  Univ of Maine - Darling Marine Cnt 
Eric Leaders Singletary  UTPB 
Laura King Sirot  Cornell University 
Owen E Sitton  Element Markets LLC 
Michael Ian Sitvarin   
Nancy Sloan  Florida Institute of Technology 
Amy Slott   
Andrew M Smith  Ithaca College 
Julian P. S. Smith  Winthrop University 
Melanie Smith  North Carolina State Univ 
Jake Socha  Virginia Tech 
Inna Sokolova  Univ of North Carolina at Charlotte 
Diara Spain  Dominican U of CA 
Trisha Spears  Florida State Univ 
Daniel Speiser  Duke University 
Heather Spence  George Washington University 
Andrew Spence  Royal Vet College 
Larry T. Spencer  Plymouth State University 
Erik Sperling  Yale University 
Simon Sponberg  UC Berkeley 
Jonathan Sprague  U of Montana 
Stephen E. Stancyk  Department of Biological Sciences University of South Carolina 
Theodore Stankowich  Univ Mass Amherst 
Daniel Stanton  Winthrop Univ 
Tiffany Stephens  Friday Harbor Labs 
Wolfgang E. Sterrer  Bermuda Natural History Museum 
Ben Stranges  Bowdoin College 
Richard R. Strathmann  University of Washington 
James Strother  Univ of California Irvine 
Torsten Hugo Struck  University of Osnabrueck 
Jennifer Stygles   
Michelle Sutton  USC Beaufort 
Billie J. Swalla  U of Washington 
Brian A Swartz  Univ of California, Berkeley 
Alison Sweeney  UC Santa Barbarba 
Marilyn F. Switzer-Dunlap  University of Hawaii, Manoa Pacific Biomedical Research Center 
Anna Syme  UCSB 
John Andrew Szczepanski  University of Rhode Island 
Alina M. Szmant  UNC 
Greg Szulgit  Hiram College 
Brett G Szymik  Eastern Connecticutt State Univ 
Kimberly Takagi  University of the Ryukyus 
David W. Tapley  Salem State College 
Sharon Tatem  Old Dominion Univ 
Maurice J. Tauber   
Carolina Tavares  Museu Nacional 
Jennifer Taylor  University of California Berkeley 
Barbara Taylor  Univ of Georgia 
Cynthia Tedore  Duke University 
Michael Temkin  St Lawrence Univ 
Jonathan Steed Terblanche  Univ of Stellenbosch 
Nora B. Terwilliger  Univ of Oregon OIMB 
Robert W Thacker  U Alabama at Birmingham 
Jamie Theobald  University California, LA 
Jana Thoma  UL Lafayette 
Kara Sue Thomas  Colorado State University 
Joseph T. Thompson  Franklin & Marshall College 
Daniel Thornhill  Bowdoin College 
Carl Thurman  University of Northern Iowa 
William J. Tiffany   
Andrea Tilden  Colby College 
Oive Tinn  UC Santa Barbara 
Cawa Tran  Univ of Hawaii 
James Traniello  Boston University 
Nikki Traylor-Knowles  Boston Univ 
Terry M. Trier  Grand Valley State University 
John R. Tripp  Florida Southern College 
Lloyd Trueblood   
Marlene Tsie  U of Maine 
Brian K. Tsukimura  California State Univ Fresno 
Shea Richard Tuberty  Appalachian State University 
Diane Tulipani  San Diego 
Richard L. Turner  Florida Institute of Technology 
Teresa Turner  Univ Virgin Islands 
Betty M. Twarog   
Kenyon S. Tweedell  University of Notre Dame Department of Biological Sciences 
Seth Tyler  University of Maine 
Catherine R.C. Unabia  Hawaii Pacific University 
Ted Uyeno  NORTHERN ARIZONA UNIVERSITY 
Linnea van Griethuijsen  Tufts University 
Leigh M. Van Valen  University of Chicago Department of Ecology & Evolution 
John Vanden Books  Arizona State University 
Thomas L. Vandergon  Pepperdine University 
Kimberly Vann  Arizona State Univ 
Dawn Vaughn  U of Washington 
Rebecca Vega  Stanford 
Jann Vendetti  UC-Berkeley 
Edward Venit  Duke University 
Cynthia Venn  Bloomsburg Univ 
Judith M. Venuti  LSU Health Science Center 
E Alan Verde  Maine Maritime Academy 
Steven Vogel  Duke University 
Kristen Vollrath   
Janice Voltzow  Univ of Scranton 
Yasmin von Dassow  Carnegie Museum of Natural History 
Michelangelo von Dassow   
Stephen A. Wainwright  Duke University 
Lindsay Waldrop  UC Berkeley 
Kelly Walsh  SFSU 
Keith Walters  Coastal Carolina University 
Linda J Walters  Univ of Central Florida 
Shi Wang  UT (Austin) 
Andreas Wanninger  University of Copenhagen 
J. Evan Ward  University of Connecticut 
Nancy E. Warner   
Les Watling  University of Hawaii at Manoa 
Kerry Weber   
Mark Webster  Univ of Chicago 
Ann M. Weigl   
Judith Shulman Weis  Rutgers University 
Marc Weissburg  Georgia Ins of Tech 
James M. Welch  Wittenberg University 
Sefton R. Wellings   
Shannon Wells  Old Dominion University 
Adrian M. Wenner  UCSB 
Timothy Werner   
Ronald R. West  Kansas State University 
Erica Westerman   
Kristen Whalen  Centre for Marine Bio-Innovation 
A. Quinton White  Jacksonville University 
Elizabeth Whitehill  Clemson University 
Lisa Whitenack  University of South Florida 
Tracy Whitford  Stroudsburg University 
Heather R Wilkinson  OXY 
Robert B. Willey  Rocky Mountain Biol. Laboratory 
Judith Williams  U of S Mississippi 
Jason Williams  UNLV 
Jason J Williams  SUNY ESF 
Heidi R Williams  Charleston Coll 
Kristina Williams   
A. O. Dennis Willows  Friday Harbor Laboratories 
Nerida Wilson  Scripps Institution of Oceanography, UCSD 
Noah Wilson-Rich  Tufts University 
Colleen Jean Winstead  Univ of Minnesota 
Judith E. Winston  Virginia Museum 
Maria Wise  Duke Marine Lab 
Gertraude Wittig   
Donna L. Wolcott  NC State Univ 
Thomas G. Wolcott  North Carolina State University Department of MEAS 
Allan F. Wolfe  Lebanon Valley Col 
Sarah A. Woodin  Univ of SC 
Gert Worheide  LMU Munich 
Mary L. Wright  UC Berkeley 
Kerri Wrinn  Miami University 
Janie Wulff  Florida State Univ 
Russell Wyeth  St Francis Xavier University 
Charles R. Wyttenbach   
Jiawu Xu  Brigham Young Univ 
David D. Yager  University of Maryland 
Steve Yanoviak  Univ of Arkansas at Little Rock 
Mona Yates  Medgar Evers 
Jeanntte Yen  Georgia Institute of Technology 
Janice E. Young  McHenry County College 
Hugh P. Young  The Steritech Group 
Jr-Kai Yu  Institute of Cellular & Organismic Biology 
Pauline Yu  Univ of Southern California 
Louis Zachos  Nat Museum of Natural Hist 
Christina Zakas  Fred C Davison Life Science Complex 
Edward J. Zalisko  Blackburn College 
William E. Zamer  Ntnl Science Fndtn 
John D. Zardus  The Citadel 
Sarah Zehr  Duke Lemur Center 
Kevin Zelnio  The Pennsylvania State Univ 
Yu Zeng  UC Berkeley 
Lingling Zhang  UT (Austin) 
Kirk Zigler  Sewanee 
Christian Zmasek